Web-building orb-weaving spiders are key generalist predators that regulate insect populations and adapt their behaviours to environmental conditions. This study investigates the web-building behaviour, architecture, and ecological role of Eriovixia excelsa in citrus plantations of Morshi and Zygiella indica in coffee plantations of Chikhaldara, Maharashtra, India. Web characteristics, including radii, sticky spirals, mesh width, hub asymmetry, capture area, and prey composition, were recorded directly in the field. E. excelsa constructed new webs daily in shaded areas between trees, with webs averaging 135–153 cm above ground and a construction time of 34.28 ± 0.25 min. Webs were symmetric (hub asymmetry index 0.10 ± 0.003), with an average mesh width of 2.48 ± 0.05 mm and capture area of 284.08 ± 6.25 cm². Z. indica repaired and reconstructed webs sequentially, including hub and spiral restoration, with total construction time of 41.58 ± 0.41 min and hub reconstruction taking 28.91 ± 1.26 min. Webs were 130–142 cm above ground, slightly asymmetric (hub asymmetry 0.34 ± 0.009), with mesh width 2.34 ± 0.03 mm and capture area 339.43 ± 9.3 cm². Prey captured by both species was dominated by Diptera, followed by Homoptera, Hymenoptera, and Coleoptera. Comparative analysis shows that E. excelsa favors routine hub-centered web construction, while Z. indica emphasizes repair and sequential assembly. These findings highlight the adaptive significance of web architecture and behavioral strategies.